Orchidaceae

Orchid Family

Jeffrey Jeanes with comments from David Jones and David Banks.

Perennial or rarely annual herbs, monopodial or sympodial, ground-(terrestrial), tree- (epiphytic) or rock-dwelling (epilithic), sometimes saprophytic (these not usually amenable to cultivation), with creeping rhizomes, thickened rootstocks or annually renewed tubers, aerial stems often thickened at the base into a pseudobulb, adventitious aerial roots sometimes abundant, terrestrial species usually forming mycorrhizal relationships with soil-borne fungi. Leaves basal or on stems, usually alternate or spirally arranged, rarely opposite, deciduous or persistent, mostly entire, usually sheathing at base, sometimes reduced to bracts or scales, usually stalkless; stipules absent. Inflorescences terminal or lateral spikes, racemes or panicles, or flowers solitary. Flowers unisexual or bisexual, more or less irregular, usually stalked, often the stalk twisting through 180 degrees inverting the flower (resupination), usually bracteate. Perianth consisting of 6 segments, an outer whorl of sepals and an inner whorl of petals, free or variously joined, rarely completely fused into a tube. Sepals 3, similar or sometimes differentiated into a dorsal and 2 laterals. Petals 3, nearly always differentiated into a median (often termed the labellum or lip) and 2 laterals (these are often similar to the sepals). Labellum usually large and showy, often 3-lobed, frequently adorned with further lobes, appendages, teeth, calli or shiny glands, sometimes with a basal spur or sac, fixed at the base, hinged and mobile or actively motile, sometimes insect-like. Stamens usually 1 (rarely 2 or 3), completely or partly fused with 2 lateral staminodes and the style to form the column (gynostemium). Anthers usually 1, mostly 4-celled, opening by longitudinal slits; pollen granular or often aggregated into bundles (pollinia), which are shed as a unit. Pollinia usually 4, but sometimes 2, 6 or 8. Ovary inferior with one cell and many ovules attached to the walls. Stigma 3-lobed.a sterile part of the median lobe (the rostellum) is sometimes involved in pollen transfer; sometimes a portion of the rostellum is modified into a viscid disc (the viscidium) and is attached to the pollinia via 1 or 2 straps of tissue (stipes). (Pollinia, stipe, caudicle and viscidium are often shed together as a unit called a pollinarium). Fruit usually a dry capsule, opening by 3 or 6 longitudinal slits, rarely indehiscent. Seeds numerous, usually minute and dust-like rarely fleshy or sclerotic.

Nearly all orchids are pollinated by animals, usually insects (some are self-pollinating or apomictic). For pollination to occur effectively the flowers have evolved various means of advertisement and reward. The pollinator is first drawn to the flower by colour, shape, movement or scent.To ensure that the pollinator will visit other flowers of the same species it is often rewarded with food in the form of nectar, pollen or pseudopollen. Elaborate mechanisms have evolved to bring about transfer of pollinia during these visitations. There is often some degree of specificity between orchids and their pollinators.

Monadenia (Disa) bracteata is a small invasive weed from S Africa covering several thousand square kilometeres in WA and widespread in SA. More recently it has invaded several areas in Vic. It is beyond the scope of this work to attempt to describe all the orchids currently in cultivation, due to the enormous number of species, hybrids and cultivars involved. Some orchids or groups of orchids may enjoy popularity for a while, and then fade into relative obscurity. Other groups such as native Australian orchids or terrestrial orchids are sometimes grown by enthusiasts, and may be only available from specialist nurseries or orchid clubs and societies. This account covers the more commonly grown orchid genera and the more popularly grown species within each genus.

The orchids as a group are readily identified by a number of distinctive characteristics, most notably the fusion of the stamens and style producing the column. A knowledge of the structure of orchid flowers and an understanding of the terminology relating to their various components, are useful aids to identification. Many of the terms that have been coined to describe the various features of the Orchidaceae are discussed in the family description above and delineated in the accompanying line drawings.

About 850 genera of about 25 000 species (about 110 genera and 1200 species native to Australia).

ORCHID STRUCTURES AND TERMS

Orchids have few uses except as ornamental plants and in the cut flower trade. Genera used by florists include: Arachnis; Cattleya, Corsage Orchid; Cymbidium; Dendrobium; Laelia; Lycaste; Odontoglossum; Paphiopedilum, Slipper Orchid; Phalaenopsis, Moth Orchid; and Vanda. Vanilla is extracted from the immature capsules of V. planifolia which is grown as vines grown on frames, 90% produced in Madagascar. Flavourings are also extracted from Angraecum and Leptotes.A gum used in musical instruments is extracted from Geodorum.

Leaf veins generally parallel along the length of the leaf; fertile stamens 1 or 2 fused with the style to form a column; pollen in masses (pollinia); third petal greatly modified to form a lip (labellum).

CONSERVATION World wide the popularity of orchids has lead to over-zealous collectors depleting wild populations, sometimes to extinction. Every care should be taken to make sure that all orchids grown in collections are obtained by legitimate means.

This account follows the classification used in Dressler (1981), and draws heavily upon that work particularly in the formulation of the key to major groups (the Tribes); Bechtel et. al. (1981); Hawkes (1965); Jones (1990); Walters et al. (1984.). Hybrids are published in Sander's List of Orchid Hybrids.The Royal Horticultural Society is the registration authority for these names and regular supplements are published to this book.

Source: Jeanes, J. (2005). Orchidaceae. In: Spencer, R.. Horticultural Flora of South-eastern Australia. Volume 5. Flowering plants. Monocotyledons. The identification of garden and cultivated plants. University of New South Wales Press.

Hero image
kingdom Plantae
phylum   Tracheophyta
class    Magnoliopsida
superorder     Lilianae
order      Asparagales
Higher taxa
Subordinate taxa
genus        Acanthephippium Blume
genus        Acianthus R.Br.
genus        Ada Lindl.
genus        Aerangis Rchb.f.
genus        Aeranthes Lindl.
genus        Aerides Lour.
genus        Amesiella Garay
genus        Angraecum Bory.
genus        Anguloa Ruiz & Pav.
genus        Ansellia Lindl.
genus        Arpophyllum La Llave & Lex.
genus        Ascocentrum J.J.Sm.
genus        Aspasia Lindl.
genus        Baptistonia Barb.Rodr.
genus        Barkeria Knowles & Westc.
genus        Bifrenaria Lindl.
genus        Bletilla Rchb.f.
genus        Brasiliorchis
genus        Brassavola R.Br.
genus        Brassia R.Br.
genus        Broughtonia R.Br.
genus        Bulbophyllum Thouars
genus        Cadetia Gaudich.
genus        Calanthe R.Br.
genus        Callerya
genus        Camaridium
genus        Catasetum Rich.
genus        Cattleya Lindl.
genus        Caucaea
genus        Ceratostylis Blume
genus        Chiloglottis R.Br.
genus        Chysis Lindl.
genus        Coelogyne Lindl.
genus        Cryptostylis R.Br.
genus        Cuilauzina
genus        Cycnoches Lindl.
genus        Cymbidium Sw.
genus        Cynorkis Thouars
genus        Cyrtostylis R.Br.
genus        Dendrobium Sw.
species        Dendrobium bigibbum
genus        Dendrochilum Blume
genus        Diplocaulobium Kraenzl.
genus        Dipodium R.Br.
genus        Disa Bergius
genus        Diuris Sm.
genus        Dockrillia Brieger
genus        Domingoa
genus        Doritis Lindl.
genus        Dracula Luer
genus        Dryadella Luer
genus        Encyclia Hook.f.
genus        Epidendrum L.
genus        Eria Lindl.
genus        Eulophia Lindl.
genus        Galeandra Lindl.
genus        Gastrochilus D.Don
genus        Geodorum Jacks.
genus        Glossodia R.Br.
genus        Gomesa R.Br.
genus        Gongora Ruiz. & Pav.
genus        Grammangis Rchb.f.
genus        Grammatophyllum Blume
genus        Grobya Lindl.
genus        Guarianthe
genus        Habenaria Willd.
genus        Huntleya Lindl.
genus        Ionopsis Humb., Bonpl. & Kunth
genus        Isabelia Barb.Rodr.
genus        Isochilus R.Br.
genus        Laelia Lindl.
genus        Lemboglossum Halb.
genus        Leptoceras (R.Br.) Lindl.
genus        Leptotes Lindl.
genus        Liparis Rich.
genus        Ludisia A.Rich.
genus        Lycaste Lindl.
genus        Masdevallia Ruiz & Pav.
genus        Maxillaria Ruiz & Pav.
genus        Miltonia Lindl.
genus        Miltoniopsis God.-Leb.
genus        Mormodes Lindl.
genus        Nageliella L.O.Williams
genus        Nanodes Rchb.f.
genus        Neofinetia Hu
genus        Neolauchea Kraenzl.
genus        Oncidium Sw.
genus        Osmoglossum Schltr.
genus        Paphiopedilum Pfitzer
genus        Peristeria Hook.
genus        Phaius Lour
genus        Phalaenopsis Blume
genus        Pholidota Hook.
genus        Phragmipedium (Pfitzer) Rolfe
genus        Physosiphon Lindl.
genus        Pinalia
genus        Plectorrhiza Dockrill
genus        Pleione D.Don
genus        Pleurothallis R.Br.
genus        Polystachya Hook.
genus        Prasophyllum R.Br.
genus        Promenaea Lindl.
genus        Psychopsis Raf.
genus        Pterostylis R.Br.
genus        Restrepia Humb., Bonpl. & Kunth
genus        Restrepiella Garay & Dunst.
genus        Rhinerrhiza Rupp
genus        Rhyncholaelia Schltr.
genus        Rhynchostele
genus        Rhynchostylis Blume
genus        Rossioglossum Garay & G.C.Kenn.
genus        Sarcochilus R.Br.
genus        Sophronitis Lindl.
genus        Spathoglottis Blume
genus        Specklinia
genus        Spiranthes Rich.
genus        Stanhopea Hook.
genus        Stelis Sw.
genus        Stenoglottis Lindl.
genus        Thunia Rchb.f.
genus        Trichocentrum
genus        Trichopilia Lindl.
genus        Vanda R.Br.
genus        Vitekorchis
genus        Zygopetalum Hook.