Mint and Sage Family

Herbs or occasionally shrubs and trees, usually with glandular hairs containing aromatic oils, and with stems that are often square in cross-section. Nonglandular hairs multicellular, uniseriate. Leaves usually opposite and generally decussate or occasionally whorled, simple to pinnately or palmately divided, usually toothed; stipules absent. Inflorescence with indeterminate main axis and determinate (cymose branched) lateral axes. Flower clusters cymose, sometimes in a head, often axillary or in the axils of leafy bracts, whorl-like (verticillaster) or sometimes solitary. Flowers bisexual, mostly irregular. Nectar disc often present. Sepals usually 5, united, persistent in fruit. Corolla usually of 5 petals united into a mostly 2-lipped tube (usually more strongly so than in Verbenaceae). Stamens 4, in 2 pairs, fused to corolla tube, sometimes only 2 fertile. Ovary superior, unlobed to deeply 4-lobed. Carpels 2, united, 2-locular, but appearing 4-locular due to development of false septa, 1 ovule per locule, with axile placentation; ovules attached to the sides of false septa. Style usually apically forked with an inconspicuous stigmatic region, terminal to arising from the central depression of 4 ovary lobes (gynobasic style). Fruit a drupe with 1–4 pits, an indehiscent, 4-seeded pod, or a schizocarp splitting into 4 nutlets or drupelets.

Although traditionally referred to as the Labiatae, the name Lamiaceae (with the standard -aceae family suffix) is now widely accepted. Phylogenetic relationships within Lamiales support a broad circumscription of Lamiaceae, including nearly two-thirds of the genera usually placed in Verbenaceae to make Lamiaceae monophyletic (Judd et al 2016). This treatment follows Stevens, P.F. (2018, Angiosperm Phylogeny Website).

The 2-lipped flowers are adapted to pollination by birds and bees, although other insects such as moths and butterflies also act as pollinators.

Many members of this family are grown as culinary herbs or, occasionally, as ornamental border plants. Tropical plants like Coleus and Plectranthus are often grown as indoor plants for the ornamental foliage.

Rarely cultivated plants include: Acinos alpinus (L.) Moench, grown as a spreading rockery plant from montane Europe; Elsholtzia stauntonii Benth., Mint Shrub, from N China, an annual subshrub to 1 m or so tall with purplish flowers; Chloanthes stoechadis R. Br. from WA, Qld and NSW, occasionally grown in warmer districts - it is a shrub to about 80 cm tall with linear leaves, greenish yellow to bluish tubular flowers, and it has often been placed in the family Chloanthaceae; Clinopodium vulgare L., Wild Basil, which has naturalised in moist forest in Victoria and flowers in summer; Dracocephalum spp., Dragonhead, from Eurasia, Africa and N America, several species of which are grown in cool-climate gardens; Faradaya splendida F. Muell., a Qld rainforest climber with shiny ovate leaves to 30 cm long, and white, fragrant, 4-lobed flowers - it is occasionally grown in warmer districts; Hemigenia purpurea R. Br., a small shrub from NSW with short linear-cylindrical leaves and purplish flowers; Horminum pyrenaicum L., Dragon Mouth, a low-growing violet-flowered rock garden plant from montane Europe; Hyptis emoryi Torr., Desert Lavender, an aromatic shrub from SW America with violet flowers; Leonurus cardiaca L., a shrub 1-2 m tall from the Mediterranean region with white or pale spotted flowers, potentially weedy but occasionally grown in herb gardens for its medicinal properties and used mainly as a tonic; Lepechinia calycina (Benth.) Epl., Pitcher Sage, a shrub to over 1m tall from California with white or pink flowers; Lycopus australis R. Br., Australian Gypsywort, from S and E Aust and L. europaeus L.; Mallophora globiflora Endl. from SW WA, a dwarf shrub with abundant, whitish, woolly-looking, globular flower heads to about 3 cm wide - it is sometimes placed in the Dicrastylidiaceae or Chloanthaceae, and Perovskia atriplicifolia Benth. from Afghanistan and Pakistan, which is occasionally grown in the border as a subshrub to about 1.5 m tall, with linear upper leaves, coarsely toothed lower leaves and spires of soft blue flowers; it is available as P. 'Blue Spire'.

Many species are culinary herbs (including Melissa, Lemon Balm; Mentha, Mint; Lavandula, Lavender; Origanum, Marjoram; Rosmarinus, Rosemary; Salvia, Sage; Thymus, Thyme; and many others).Timber is obtained from Tectona,Teak; and Vitex. In the tropics Pogostemon is the source of patchouli oil and Perilla is grown in India for perilla oil, which is used in paints and printing inks. Indigenous peoples have many uses for members of this family: Sideritis is used to make a tea and is sometimes available commercially as dried foliage and flower heads; and tubers of Coleus rotundifolius are edible and similar to the potato.

For differences from Verbenaceae, see the discussion for the latter.

About 236 genera and c. 7200 species, cosmopolitan, found in most habitats and altitude ranges but with a centre of distribution around the Mediterranean. Australia has over 40 genera.

Harley & Reynolds (1992).

Source: Spencer, R. (2002). Lamiaceae. In: Spencer, R.. Horticultural Flora of South-eastern Australia. Volume 4. Flowering plants. Dicotyledons. Part 3. The identification of garden and cultivated plants. University of New South Wales Press.

Updated by: Val Stajsic, April 2018

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kingdom Plantae
phylum   Tracheophyta
class    Magnoliopsida
superorder     Asteranae
order      Lamiales
Higher taxa
Subordinate taxa
genus        Agastache Gronov
genus        Ajuga L.
genus        Ballota L.
genus        Bystropogon L'Hér.
genus        Calamintha Mill.
genus        Callicarpa L.
genus        Caryopteris Bunge
genus        Cedronella Moench.
genus        Clerodendrum L.
genus        Glechoma L.
genus        Gmelina L.
genus        Hemiandra R.Br.
genus        Holmskioldia Retz
genus        Hyssopus L.
genus        Lamium L.
genus        Lavandula L.
genus        Leonotis (Pers.) R.Br.
genus        Marrubium L.
genus        Melissa L.
genus        Mentha L.
genus        Moluccella L.
genus        Monarda L.
genus        Nepeta L.
genus        Ocimum L.
genus        Origanum L.
genus        Phlomis L.
genus        Physostegia Benth.
genus        Plectranthus L'Hér.
genus        Premna L.
genus        Prostanthera Labill.
genus        Prunella L.
genus        Rosmarinus L.
genus        Rotheca
genus        Salvia L.
genus        Satureja L.
genus        Solenostemon Thonn.
genus        Stachys L.
genus        Teucrium L.
genus        Thymus L.
species        Thymus praecox
genus        Vitex L.
genus        Westringia Sm.