Trees or shrubs, lacking latex, deciduous or not, monoecious or dioecious. Leaves usually alternate (rarely opposite or whorled), simple to palmate, marings entire to rarely spinose-dentate, petiolate; stipules present or absent, often caducous. Inflorescences axillary, cymose, racemose, spicate or paniculate, bracteate. Flowers regular, unisexual, pedicellate; sepals 3–14 sepals , free; petals absent. Male flowers with 3–14 usually free sepals and (2–)4–10(–55) stamens, their filaments free or connate, anthers dehiscing by longitudinal slits. Female flowers with as many sepals as males, ovary superior, (2–)3(–5) locular, ovules (1–)2 per locule, nectary disc present or absent below ovary, annular or lobed; styles (2–)3–5, distinct,  undivided or bifid. Fruit a septicidally dehiscent schizocarp, or rarely a drupe. Seeds 1 or 2 per locule, often carunculate.

About 24 genera, and about 96 species, predominantly from S. Hemisphere (Africa, Madagascar, Central and South America, tropical Asia, New Guinea, Australia, New Caledonia) but also in SW United States; 9 genera in Australia and about 42 species.

Formerly included as a subfamily (Oldfieldioideae) within the Euphorbiaceae from which it differs in absence of latex and ovaries with 2-ovules per locule. This family has long been recognized as monophyletic because of its distinctive spiny pollen.

Created by: Val Stajsic

Updated by: Val Stajsic, April 2018

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kingdom Plantae
phylum   Tracheophyta
class    Magnoliopsida
superorder     Rosanae
order      Malpighiales
Higher taxa
Subordinate taxa
genus        Pseudanthus Sprengel